In April 1945, the Western Allied invasion of Germany encountered strong resistance. Don “Wardaddy” Collier, a battle-hardened staff sergeant in the U.S. Second Armored Division, commands an M4 Sherman nicknamed Fury whose crew consisting of gunner Boyd “Bible” Swan, loader Grady “Coon-Ass” Travis, driver Trini “Gordo” Garcia, and assistant driver–bow gunner “Red” have been together since the North African campaign.

After Red is killed in a tank battle, he is replaced by Private First Class Norman Ellison, a young typist clerk from V Corps. As Fury moves deeper into Germany, the crew disdains Norman’s combat inexperience and aversion to violence. While moving in a tank convoy, he spots but does not fire upon a group of concealed Hitler Youth child soldiers, who promptly ambush the tank column with a Panzerfaust, destroying the lead tank and killing the platoon leader and his crew. Don assumes command of the tank column. Later, while in a battle, Norman hesitates under fire from anti-tank guns.

After the battle, Don orders Norman to execute a captured German soldier. When he refuses, Don wrestles his revolver into Norman’s hand and forces him to pull the trigger, killing the soldier and traumatizing Norman. After the platoon captures the small town of Kirchohsen, Don takes Norman to help search and clear an apartment, where they discover two German women – Irma and her younger cousin Emma – in hiding. After Don gives them some supplies, the women prepare a hot meal and give Don hot water for a shave.

As they begin to bond, Norman and Emma retreat to the bedroom and have sex, having been prompted to do so by Don. Later, as the four sit down to eat together, the rest of Fury’s crew drunkenly barges in, harassing the women and bullying Norman. The crew’s resentment towards Norman leads to a tense standoff with Don before they all are called away for an urgent mission. As they depart, German artillery strikes the town and kills Emma, causing Norman to suffer a mental breakdown. The four tank crews receive orders to hold a vital crossroads to protect the division’s vulnerable rear lines.

En route, a lone German Tiger I ambushes and destroys the other three tanks before being defeated by Fury. The radio gets damaged in the battle, preventing Don from calling for reinforcements. Don decides that he and his crew must continue the mission alone. When they arrive at the crossroads, Fury is immobilized by a landmine. While the tank is being repaired, Don sends Norman to scout a nearby hill, where he spots a battalion of Waffen-SS marching towards Fury’s position. When Norman returns to inform the crew, all but Don plan to retreat. Refusing to leave, Don insists he will stay to hold the crossroads and tells the others to leave. After Norman elects to stay with Don, the others ultimately stay on as well.

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